Thirty employees working at the Paris landmark before the fire broke out were interviewed by Paris prosecutors on Tuesday, the Paris prosecutor's office said. Those were due to continue on Wednesday.
The Paris fire service, meanwhile, said the nine-hour battle to save the building was one of them most complex it had ever undertaken. At one point, it was feared that the entire structure might be lost
"If the flames had actually got to the timber frames of the belfries then we would have lost the cathedral completely because it would have led to a chain reaction of collapse," said Gabriel Plus, a spokesman for the fire service.
Philippe Demay, Deputy Chief of the Paris Fire Brigade, told reporters on Wednesday that the operation to put out the flames was "complex," and the "most complicated" he had ever encountered.
On Wednesday evening, cathedrals across France rang their bells in honor of Notre Dame, marking two days since the fire. Bells tolled at 6:50 p.m. across the country acknowledging what the French Bishops Conference described as "a shock that affects far beyond just the Catholics of our country."
The French government said it was committed to rebuilding the the cathedral. Prime Minister Edouard Philippe announced an international architects' competition to replace the building's fallen spire.
As the scale of the damage became clear, investigators said they were determined to get to the bottom of how the blaze started. "Investigations continue to search for the truth and identify the origins and causes of the fire," the Paris prosecutor's office said.
Scrutiny has fallen on the firms undertaking the renovation of the 150-year-old spire, which collapsed Monday as the flames raged around it.
Of the four companies contracted to carry out renovations at Notre Dame, only two companies, scaffolding firm Europe Echafaudage and art conservationists Socra, had work in progress there at the time of the fire. Neither company had workers on site when the fire broke out, CNN has been told.
According to centraledesmarches.com, a website where private companies can bid for public projects, Europe Echafaudage and its parent company, Le Bras Freres, were awarded contracts worth 3,493,766 euros ($3,951,396) for scaffolding and other services.
Julien Le Bras, CEO of Europe Echafaudage, which received a contract to renovate the spire in 2017, told reporters Tuesday that 12 of his employees were working on the project, but that "there were absolutely no workers on site when the fire broke out, and there hadn't been for quite a while before the fire."
Mark Eskenazi, a PR representative for Le Bras Freres, told CNN the company is speaking to experts about how to take down the scaffolding and "absolutely" denies responsibility for what happened, saying that its workers had left the cathedral one hour before the fire began. He called the fire a "crisis" for the small business of 20 workers. "They are artisans, it is a very small enterprise."
Patrick Palen, a spokesman for Socra, which was brought in to refurbish statuettes from the cathedral, told CNN that workers from the company removed 12 statues of the apostles and four of evangelists from the cathedral last week. Those statues were being held safely in an off-site workshop at the time of the fire. Palen said Socra employees were not at Notre Dame during the fire.
Questions have been raised about why the blaze was not tackled sooner. Security guards at the 850-year-old cathedral first heard the fire alarm at 6:20 p.m. local time on Monday, and evacuated the cathedral even though they didn't see any sign of fire, a spokesman for the Paris Fire Brigade told CNN.
The alarm sounded again some 20 minutes later, at which point the cathedral's security officers noticed the fire, Paris prosecutor Remy Heitz said.
Fire chiefs denied that they were slow to react to the initial reports of the blaze.
Despite the intensity of the fire, authorities have said that some of the cathedral's major relics, artifacts and artwork had been rescued. Some of the most valuable artworks are being held at City Hall, while a number of other pieces are being transferred to the Louvre museum.
The Crown of Thorns, believed by many to be a relic of the passion of Christ, and the Tunic of Saint Louis were saved. A copper rooster that sat atop the spire which collapsed in Monday's blaze has been recovered intact.
Le Parisien newspaper, citing a source from the Ministry of Culture, said it was not yet known if relics contained in the rooster had survived the blaze.
Donations towards the restoration have been rising steadily; the total raised so far for the reconstruction of the Gothic masterpiece has topped 800 million euros ($904 million), including gifts of tens of millions of euros from some of France's wealthiest families.
French billionaire Bernard Arnault, chairman and CEO of luxury good consortium LVMH, pledged $200 million, adding to a purse of $55 million promised by the city of Paris.
Cosmetics company L'Oréal, along with The Bettencourt Meyers family and the Bettencourt Schueller Foundation, has donated 200 million euros ($226 million) to help fund the repairs.
The Vatican said it will offer technical know-how on rebuilding the cathedral. "We have the type of know-how that the whole world recognizes as being of high quality," Cardinal Gianfranco Ravasi told reporters. "So I think an eventual future offer by the Holy See will be significant."
Prime Minister Edouard Philippe said the contest to replace the spire "will be organised in due course." A committee will be set up to oversee the project and will decide whether the spire should be reconstructed or "adapted to modernity," he said.
Philippe did not say if the rebuild of the cathedral would be finished in time for the 2024 Paris Olympics, but said the work would be done as fast as possible.
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